That's quite a different shade of FG and I definitely like it.
As for the reverse head white 4001, note the sealed tuners aswell. Maybe it's a custom order indeed.
The 1999 looks just like a C64 with a proper oriented headstock. And the pic in the japanese site is a photoshoped version of the C64 pic.
The new 1999 isn't that much different from a v63, which can be had for a lot less than the RRP of the Japan LE.
The only real difference that I can see (other than the tug bar) is the new (and attractive) Fireglo. Some FG v63's have the 90's clown burst.
The V63 has a lot of differences compared to the C64: different head shape, different body - most noticeable on the horns, but also on the edge countours-, different pickguard shape and size, different horse-shoe mounting ring, different bridge, difference in the dot markers...
Not to be picky, but I just wanted to point that out...
From the factory pics, body shape of those 1999 looks just like C64, but the head looks more like the newer 4003 heads rather than the C64.