Lennon's woody, zero-sustain and badly intonated 325 is like the real rhythm keeper on those early Beatles cuts! It kept right out the way of George's lead lines and drove the songs along. Not a tone I would want to be getting, but quite recognisable.
They did have a clear good tone / bad tone strategy, but personally when choosing a guitar I'm more inclined to go for the good tone shelf rather than the so-bad-it's-immediately-recognizable. But to each their own
I will give them this — when Paul came back from the store with his new Epi Casino, John and George immediately ordered theirs. And it did not compromise their performance. John could still get his signature crappy tone, and George, you know, could make anyhing sound good.